When a young Edwardian family leaves the shores of England to build a home in the wilderness of East Africa, what they encounter is beyond their imagination, but forever remembered through the eyes of their 11-year-old daughter. Based on the beloved memoir by Elspeth Huxley, The Flame Trees of Thika brings to life the colour and adventure of turn-of-the-century Kenya. In 1913, Robin (David Robb) and Tilly Grant (Hayley Mills) arrive in Kenya with the dream of transforming a barren plot of land into a thriving coffee plantation. But torrential rains, relentless insects and murderous animals, as well as relations with natives and other settlers, challenge their ambitions. Will England call them back? Or will the wide-eyed wonder of young Elspeth (Holly Aird) help unlock the mysteries of a foreign land and open the doors into the pleasures and rewards of a new home?
Young Elspeth arrives in Thika, Kenya, with her optimistic parents who have purchased an unpromising tract of land in hopes of establishing a coffee farm.
The Grants get new neighbors when Hereward and Lettice Palmer arrive. He was an officer in the British Indian Army and neither seem prepared for the hardship they must face. Lettice almost goes to pieces when she finds their headman has been badly beaten and lay dying. She tells her husband that that she's not sure she wants to stay. Mr. Roos offers to show Robin's workers how to handle the plow animals but Robin doesn't quite agree with his handling of the local Africans, including Sammy. Elspeth receives a gift of two chameleons which she names George and Mary.
At the end of 1913, Tilly and Robin travel to Nairobi with the Palmers to ring in the new year. They leave Eslpeth in the care of Mrs. Nimmo, whose husband is still away. On New Year's eve, Mrs. Nimmo has a party and invites Alec Wilson and another couple. A good time is had by all - even when one of the guests begins to shoot up the place. A friend of the Palmers, Ian Crawford, arrives after acquiring horses in Abyssinia and Elspeth receives a very special gift. Crawford is very much attracted to Lettice but she resists his advances.
Elspeth and Sammy are in put in danger when they face a herd of rampaging elephants. Mrs. Nimmo attends to Sammy's wife who has had a miscarriage, but her recommendation that the woman be transported to the hospital in Nairobi is overruled when the family brings in the local medicine man. Robin has been blasting old trees and stumps, but tragedy ensues when Sammy's young son steals blasting caps from a locked shed. Njombo becomes mysteriously ill and while the doctor can find nothing wrong with him, he is convinced that a curse has been placed on him, most likely by Sammy who seems to blame him for his recent losses.
The Palmers receive their new piano and face a challenge in getting it moved to the farm. Hereward is worried about Lettice and whether she has the stamina to bear living in Africa. He notes that about the only activity she engages in is to go on safari with Ian Crawford. As Lettice and Hereward's marriage begins to disintegrate, it is apparent to almost everyone that Ian and Lettice are in love. When one of Lettice's beloved pekingese is carried away by a leopard, Hereward, Robin and Alec Wilson set off to kill it. They eventually turn to Mr. Roos for some professional advice.
With Robin and Elspeth staying behind at the farm, Tilly joins the Palmers and Ian Crawford on safari. Lettice finally admits to Ian that she loves him. Things come to a boil between Ian and Hereward but Crawford's bearer Ahmed intervenes, severely wounding Palmer. At the farm meanwhile, Elspeth takes Njomo's advice and gets the local medicine man to give her a charm to protect Twinkle. She's hurt however when Twinkle goes off on her own and doesn't return home. Everyone hears that World War I has started in Europe and Robin and the Major decide to go to Nairobi see what is happening.
With Robin away, Tilly decides to go to Nairobi to do what she can for he war effort. As a result, Elspeth must go lives with the Robinsons, a wealthy merchant family who have a resident governess for their children. Elspeth doesn't adapt very well however and decides to run away. Hereward is in the army in France while Robin and Ian Crawford have joined the local forces but Tilly gets some bad news after a battle in the south. When Robin gets orders to join his old regiment in France, Tilley make plans to return to England, at least temporarily.
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